diff --git a/lib/galaxy/tools/parameters/basic.py b/lib/galaxy/tools/parameters/basic.py
index 34b325a0d123..51244d5dd927 100644
--- a/lib/galaxy/tools/parameters/basic.py
+++ b/lib/galaxy/tools/parameters/basic.py
@@ -242,6 +242,9 @@ def to_json(self, value, app, use_security) -> str:
 
     def to_python(self, value, app):
         """Convert a value created with to_json back to an object representation"""
+        if value is None and (default_value := getattr(self, "value", None)) not in ("", None):
+            # None should be equivalent to no value provided, so use default
+            return default_value
         return value
 
     def value_to_basic(self, value, app, use_security=False):
@@ -494,6 +497,9 @@ def to_python(self, value, app):
         except (TypeError, ValueError) as err:
             if contains_workflow_parameter(value):
                 return value
+            if value is None and self.value not in ("", None):
+                # None should be equivalent to no value provided, so use default
+                return int(self.value)
             if not value and self.optional:
                 return None
             raise err
@@ -567,6 +573,9 @@ def to_python(self, value, app):
         except (TypeError, ValueError) as err:
             if contains_workflow_parameter(value):
                 return value
+            if value is None and self.value not in ("", None):
+                # None should be equivalent to no value provided, so use default
+                return float(self.value)
             if not value and self.optional:
                 return None
             raise err
@@ -623,6 +632,9 @@ def from_json(self, value, trans, other_values=None):
         return self.to_python(value)
 
     def to_python(self, value, app=None):
+        if value is None and self.checked is not None:
+            # None should be equivalent to no value provided, so use default
+            return self.checked
         if not self.optional:
             ret_val = string_as_bool(value)
         else: