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That would be the expand tags stepping up the tree (reference subject role -> participant under investigation role -> role, Ontobee) (code responsible).
If it's not being dropped it's because there's (at least) one experiment annotated with "role" and not "reference subject role", otherwise no new unique chain would be found (code) and it would be filtered. I guess it's also possible that it expanded from something outside of Uberon too since "role" is used in quite a few ontologies (Ontobee).
Should be able to track this down by just grepling ("role") in cf.Base and cf.Val (or whatever they're called).
Something splits reference subject role and just "role" into two separate tags
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